The Fishes of the Inland Waters of Southeast Asia : A Catalogue and Core Bibliography of the Fishes Known to Occur in Freshwaters, Mangroves and Estuaries, Maurice Kottelat.
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Family BADIDAE
Badidae Barlow, Liem & Wickler, 1968
Badidae Barlow, Liem & Wickler, 1968: 444 (type genus:
Badis Bleeker, 1853o: 106)
Badis Bleeker, 1853
Badis Bleeker, 1853o: 106 (type species: Labrus badis
Hamilton, 1822: 70, by absolute tautonymy). Gender
masculine.
Badis corycaeus Kullander & Britz, 2002
Badis corycaeus Kullander & Britz, 2002: 342, fig. 26 (type
locality: Myanmar: Kachin State: Myitkyina district: Irrawaddy
River drainage, Nan Kywe Chaung about 200
m South of Sha Dau village, about 18 km on road Myitkyina�
Mogaung; 25�20'03"N 97�16'40"E; holotype:
NRM 36193)
Badis ferrarisi Kullander & Britz, 2002
Badis ferrarisi Kullander & Britz, 2002: 332, fig. 19 (type
locality: Myanmar: Saigaing Division: Chindwin River
drainage, Kalaymyo markets: reported to come from
Myit-tha River and nearby hillstreams of Myit-tha drainage;
holotype: NRM 48424)
Badis juergenschmidti Schindler & Linke, 2010
Badis juergenschmidti Schindler & Linke, 2010: 210,
figs. 2�5 (type locality: Myanmar: Ka Dat Chaung River,
at Kammon Chaung village, about 8 km northwest of
Kyaiktiyo, 17�24'22"N 97�04'20"E; holotype: MTD
F 32325)
Badis khwae Kullander & Britz, 2002
Badis khwae Kullander & Britz, 2002: 328, fig. 18 (type
locality: Thailand: Kanchanaburi Province: small stream
near Ban Chong Lu, about 13 km northwest of Sangkhlaburi;
15�10'21"N 98�20'00"E; holotype: NRM 47959)
Badis kyar Kullander & Britz, 2002
Badis kyar Kullander & Britz, 2002: 350, fig. 30 (type locality:
Myanmar: Kachin State: Myitkyina district: Irrawaddy
River drainage, Nan Kywe Chaung about 200 m
south of Sha Dau village, about 18 km on road Myitkyina�
Mogaung; 25�20'3"N 97�16'40"E; holotype: NRM
37302)
Badis pyema Kullander & Britz, 2002
Badis pyema Kullander & Britz, 2002: 346, fig. 28 (type
locality: Myanmar: Kachin State: Irrawaddy River drainage,
Nan Hto Chaung, in Putao, about 1 mile from 46th
regiment; 27�19'44"N 97�22'36"E; holotype: NRM
43005)
Badis ruber Schreitm�ller, 1923
Badis badis var. rubra Schreitm�ller, 1923: 254 (type locality:
Myanmar: Bago Division: Irrawaddy River drainage,
roadside ditch, about 48 km on road Pyay-Yangon,
18�30'41"N 95�24'48"E [original locality: reportedly
India (aquarium material)]; neotype: NRM 36194, designated
by Kullander & Britz, 2002: 325, fig. 13a)
Badis badis burmanicus Ahl, in Arnold & Ahl, 1936: 449,
fig. (type locality: Burma: Rangoon; syntypes [3]: ZMB
31904 [1], Paepke, 1993b: 93, fig. 1; also in Ahl, 1937: 118)
Badis siamensis Klausewitz, 1957
Badis badis siamensis Klausewitz, 1957a: 199, fig. 3 (type
locality: Thailand: Phuket Island: Phuket; holotype: SMF
3978)
Dario Kullander & Britz, 2002
Dario Kullander & Britz, 2002: 354 (type species: Labrus
dario Hamilton, 1822: 72, 368, by original designation).
Gender masculine.
Dario dayingensis Kullander & Britz, 2002
Dario dayingensis Kullander & Britz, 2002: 364, fig. 39
(type locality: China: Yunnan: Ying Jiang County: Irrawaddy
River drainage: Jiu Cheng town: small stream
tributary to Da Ying Jiang River, at upstream end of Bing
Han Cun village; 24�45'50"N 98�8'54"E; holotype: NRM
21118)
Dario hysginon Kullander & Britz, 2002
Dario hysginon Kullander & Britz, 2002: 359, figs. 36�37
(type locality: Myanmar: Kachin State: Myitkyina district:
Irrawaddy River drainage: Nan Kywe Chaung under
bridge on road south to Mogaung; 25�20'20"N
97�16'57"E; holotype: NRM 44004)
Acanthophthalmus Bleeker, 1858i: 304 (incorrect subsequent
spelling of Acantophthalmus van Hasselt, 1823c: 132;
not a junior homonym of Acanthophthalmus van Hasselt,
1824b: 376 as this is also an incorrect subsequent
spelling of Acantophthalmus van Hasselt, 1823c: 132 and
on Official Index of Rejected and Invalid Generic Names
in Zoology, ICZN, 1992b: 248 [Opinion 1695]).
Pangio Blyth, 1860b: 169 (type species: Cobitis cinnamomea
M'Clelland, 1839: 304, by monotypy). Gender feminine
[Code art. 30.2.3].
Apua Blyth, 1860b: 169 (type species: Apua fusca Blyth,
1860b: 169, by monotypy; simultaneous subjective synonym
of Pangio Blyth, 1860b: 169, first reviser [Kottelat,
1987a: 371] gave precedence to Pangio). Gender feminine.
Eucirrhichthys Perugia, 1892: 1009 (type species: Eucirrhichthys
doriae Perugia, 1892: 1009, by monotypy).
Gender masculine.
Cobitophis Myers, 1927: 4 (type species: Acanthophthalmus
anguillaris Vaillant, 1902: 151, by original designation).
Gender masculine.
Pangio agma (Burridge, 1992)
? Acanthophthalmus borneensis Boulenger, 1894a: 251 (type
locality: Malaysia: Borneo: Sarawak: Baram River; syntypes:
BMNH 1889.7.31.14 [1], MNHN 1894.18 [1],
Bertin & Est�ve, 1948: 91, Eschmeyer, 2010, Kottelat et
al., 1993: pl. 29)
Acanthophthalmus agmus Burridge, 1992: 180, fig. 6 (type
locality: Malaysia: Borneo: Sarawak: Fourth Division:
Sungai Lawa, 1 km upstream of confluence with Baram
River; holotype: ROM 58726)
Pangio alcoides Kottelat & Lim, 1993
Pangio alcoides Kottelat & Lim, 1993: 208, fig. 1 (type locality:
Malaysia: Terengganu: Rantau Abang, stream at
154-km stone on road from Kuala Terengganu to Kuantan;
holotype: ZRC 17209)
Pangio alternans Kottelat & Lim, 1993
Pangio alternans Kottelat & Lim, 1993: 210, fig. 3 (type
locality: Indonesia: Borneo: Kalimantan Timur: Mahakam
River basin: blackwater stream entering Mahakam River
on northern side near Mujub; 0�01'S 115�43'E; holotype:
MZB 5895)
Pangio anguillaris (Vaillant, 1902)
Acanthophthalmus anguillaris Vaillant, 1902: 151, fig. 46
(type locality: Indonesia: Borneo: Kalimantan Barat:
Kapuas River, possibly at Sintang; holotype: RMNH
7788, Eschmeyer, 2010 [a single specimen implied
p. 153])
Acanthophthalmus vermicularis Weber & de Beaufort, 1916:
34 (type locality: Indonesia: Sumatra: Kampar Kiri River;
holotype: ZMA 100.260, Nijssen et al., 1982: 26)
Cobitophis perakensis Herre, 1940a: 8, pl. 2 (type locality:
Malaysia: Perak: lake above Chenderoh Dam; holotype:
CAS-SU 33004, B�hlke, 1953: 39)
Taxonomic notes. Several species are confused under this
name. See also molecular data in Bohlen et al. (2011).
Pangio atactos Tan & Kottelat, 2009
Pangio atactos Tan & Kottelat, 2009: 57, fig. 42 (type locality:
Indonesia: Sumatra: Jambi: Danau Kamining near
Kampung Trasos, about 5 km southwards on unpaved
road turning-off from Muara Bungo�Muara Tebo road at
km 36; holotype: MZB 10995)
Pangio bitaimac Tan & Kottelat, 2009
Pangio bitaimac Tan & Kottelat, 2009: 59, fig. 43 (type locality:
Indonesia: Sumatra: Jambi: Sungai Alai; holotype:
MZB 10997)
Pangio cuneovirgata (Raut, 1957)
Acanthophthalmus cuneovirgatus Raut, 1957: 30, figs. (type
locality: unknown, possibly "Jahore/Hinterindien" [Malaysia:
Johor]; holotype: ZMB 21332)
Pangio doriae (Perugia, 1892)
Eucirrhichthys doriae Perugia, 1892: 1009 (type locality:
Malaysia: Borneo: Sarawak; syntypes: MCSNG 9231 [4],
ZMA 114.898 [1], Tortonese, 1961: 188, Nijssen et al.,
1982: 28)
Pangio elongata Britz & Maclaine, 2007
Pangio elongata Britz & Maclaine, 2007: 18, fig. 1 (type
locality: Myanmar: Tenasserim: mouth of Mingaw Kloh
[Min Ngaw Chaung, stream entering Tenasserim River
at 13�25'N 99�01'E]; holotype: BMNH 1992.11.16.13)
Pangio filinaris Kottelat & Lim, 1993
Pangio filinaris Kottelat & Lim, 1993: 219, fig. 9 (type locality:
Malaysia: Terengganu: Sungai Tersat, a tributary
of Sungai Terengganu, immediately downriver of Sekayu
Waterfall Park, 4�57'51"N 102�57'45"E; holotype:
ZRC 34915)
Pangio fusca (Blyth, 1860)
Apua fusca Blyth, 1860b: 169 (type locality: Burma: Tenasserim;
syntypes [3]: ZSI F 2647/1 [2], ? AMS B.7500
[1], Hora, 1921d: 32, Menon & Yazdani, 1968: 120, Eschmeyer,
2010)
Pangio incognito Kottelat & Lim, 1993
Pangio incognito Kottelat & Lim, 1993: 221, fig. 11 (type
locality: Borneo: Sarawak: km 42 on road from Lindu to
Kuching, west of Sungei Stinggang; holotype: ZRC
34913)
Pangio kuhlii (Valenciennes, in Cuvier & Valenciennes,
1846)
Acantophthalmus fasciatus van Hasselt, 1823c: 133 (nomen
nudum, Kottelat, 1987a: 371)
Acanthophthalmus fasciatus van Hasselt, 1824b: 377 (nomen
nudum)
Cobitis kuhlii Valenciennes, in Cuvier & Valenciennes, 1846:
77 (type locality: Indonesia: Java: Krawang [original type
locality: Indonesia: Java: surroundings of Batavia [Jakarta]];
neotype: RMNH 2688, designated by Burridge, 1992:
182 [holotype listed by Bertin & Est�ve, 1948: 91, Eschmeyer,
2010, has no type status, see Burridge, 1992: 181])
Acanthophthalmus fasciatus Bleeker, 1860j: 74 (type locality:
Indonesia: Java: Krawang [original type locality: Indonesia:
Java: Batavia [Jakarta], Buitenzorg [Bogor],
Penawangan / Sumatra: Lahat]; neotype: RMNH 2688,
designated by Kottelat & Lim, 1993: 224; objective junior
synonym of Cobitis kuhlii Valenciennes, in Cuvier &
Valenciennes, 1846: 77)
Pangio lidi Hadiaty & Kottelat, 2009
Pangio lidi Hadiaty & Kottelat, 2009b: 65, figs. 1�2 (type
locality: Indonesia: Kalimantan Timur: Mahakam River
drainage, Ulu Belayan, Sungai Petung Kanan (0�32'06"N
116�10'55"E); holotype: MZB 16528)
Pangio longimanus Britz & Kottelat, 2010
Pangio longimanus Britz & Kottelat, 2010: 373, fig. 1 (type
locality: Laos: Vientiane Province: confluence of Nam
Leuk and Nam Gnong, 18�22'04"N 103�05'27"E; holotype:
ZRC 51933; proposed as compound noun in apposition,
indeclinable)
Pangio lumbriciformis Britz & Maclaine, 2007
Pangio lumbriciformis Britz & Maclaine, 2007: 22, fig. 4
(type locality: Myanmar: Irrawaddy basin: Nan Kwe
stream [near Myitkyna]; holotype: BMNH 2006.9.29.6)
Pangio malayana (Tweedie, 1956)
Acanthophthalmus kuhlii malayanus Tweedie, 1956: 58,
pl. 6a (type locality: Malaysia: Pahang: Kuala Tahan; holotype:
BMNH 1957.1.23.1)
Pangio mariarum (Inger & Chin, 1962)
Acanthopthalmus mariae Inger & Chin, 1962: 118, fig. 56
(type locality: Malaysia: Borneo: Sabah: Kinabatangan
District: small tributary of Kinabatangan River at Deramakot;
holotype: FMNH 68161; incorrect original spelling,
must be emended to mariarum [Code art. 31.1.2],
Kottelat, 1989: 13; Kottelat, 2012d: 39)
Pangio muraeniformis (de Beaufort, 1933)
Acanthophthalmus muraeniformis de Beaufort, 1933: 32
(type locality: Singapore: Thomson Road; syntypes: ZRC
1052 [2], ZMA 103.185 [3], CAS-SU 32602 [1], Nijssen
et al., 1993: 213, B�hlke, 1953: 39)
Pangio myersi (Harry, 1949)
Acanthophthalmus myersi Harry, 1949: 69 (type locality:
Southeastern Thailand: Nong Khor; holotype: USNM
103300)
Pangio oblonga (Valenciennes, in Cuvier & Valenciennes,
1846)
Acantophthalmus Javanicus van Hasselt, 1823c: 133 (nomen
nudum, Kottelat, 1987a: 371)
Acanthophthalmus javanicus van Hasselt, 1824b: 377 (nomen
nudum)
Cobitis oblonga Valenciennes, in Cuvier & Valenciennes,
1846: 76 (type locality: Indonesia: Java: Bogor [original
type locality: Indonesia: Java: surroundings of Buitenzorg
[Bogor]]; neotype: ZRC 35047, designated by Kottelat
& Lim, 1993: 234 [RMNH 2710 listed as possible
syntypes by Eschmeyer, 2010 cannot be syntype as Valenciennes
based his description on a drawing of a single
specimen])
Acanthophthalmus javanicus Bleeker, 1860j: 75 (type locality:
Indonesia: Java: Bogor [original type locality: Indonesia:
Java: Buitenzorg [Bogor], Tjampea / Sumatra:
Lahat]; neotype: ZRC 35047, designated by Kottelat &
Lim, 1993: 235 [RMNH 7061 listed by Eschmeyer, 2010
are not syntypes; a lectotype designation by Kottelat &
Lim, 1993: 235 made them paralectotype if they were
syntypes; the lectotype is lost and a neotype was designated];
junior objective synonym of Cobitis oblonga Valenciennes,
in Cuvier & Valenciennes, 1846: 76)
Taxonomic notes. Several species are confused under this
name. See also molecular data in Bohlen et al. (2011).
Pangio pangia (Hamilton, 1822)
Cobitis pangia Hamilton, 1822: 355, 394 (type locality: India:
"north-eastern parts of Bengal" [Goalpara; Hora,
1935: 49]; types: NT; on Official List of Specific Names
in Zoology, ICZN 1992b: 248 [Opinion 1695]; Hamilton's
unpublished figure reproduced in M'Clelland, 1839:
pl. 51 fig. 5, Britz & Maclaine, 2007: 25, fig. 6)
Cobitis cinnamomea M'Clelland, 1839: 304, 435, pl. 51 fig. 5
(unnecessary replacement name for Cobitis pangia Hamilton,
1822: 335)
Canthophrys rubiginosus Swainson, 1839: 310 (available by
indication to Hamilton, 1822: "Cob. No. 6" [which is
Cobitis pangia]; type locality: India: "north-eastern parts
of Bengal" [Goalpara; Hora, 1935: 49]; types: NT; Hamilton's
unpublished figure reproduced in M'Clelland, 1839:
pl. 51 fig. 5)
Taxonomic notes. At least two species are confused under
this name. See also molecular data in Bohlen et al. (2011).
Pangio piperata Kottelat & Lim, 1993
Pangio piperata Kottelat & Lim, 1993: 236, fig. 20 (type
locality: Malaysia: Terengganu: stream at about km 6 on
road from Kuala Brang to Kuala Terengganu, 6�04'25"N
103�03'20"E; holotype: ZRC 35003)
Taxonomic notes. At least two species are confused under
this name. See also molecular data in Bohlen et al. (2011)
Pangio pulla Kottelat & Lim, 1993
Pangio pulla Kottelat & Lim, 1993: 238, fig. 21 (type locality:
Indonesia: Kalimantan Tengah: heath forest northwest
of Palankaraya; blackwater stream, 28�29 km on
road to Tangkiling; tributary of Sungei Rungan, Sungei
Kahajan drainage; holotype: ZRC 35022)
Pangio robiginosa (Raut, 1957)
Acanthophthalmus robiginosus Raut, 1957: 31, figs. (type
locality: Indonesia: Java: stream Tjipaja-en and tributaries
near Rangkas-Betong village, Bantean area; holotype:
ZMB 21334)
Pangio semicincta (Fraser-Brunner, 1940)
Acanthophthalmus semicinctus Fraser-Brunner, 1940: 172,
fig. 3 (type locality: Malaysia: Johor: Mawai district;
holotype: BMNH 1938.12.1.113)
Acanthophthalmus kuhlii sumatranus Fraser-Brunner, 1940:
175, fig. 4B (type locality: Sumatra: Palembang Province:
Lahat; holotype: BMNH 1866.5.2.41; simultaneous
subjective synonym of Acanthophthalmus semicinctus
Fraser-Brunner, 1940: 172, first revisers [Tan & Kottelat,
2009: 61] gave precedence to A. semicinctus)
Taxonomic notes. Several species are confused under this
name. See also molecular data in Bohlen et al. (2011).
Pangio shelfordii (Popta, 1903)
Acanthophthalmus shelfordii Popta, 1903: 231, fig. (type
locality: Malaysia: Borneo: Sarawak: Sarawak River near
Kuching; holotype: RMNH 7661)
Taxonomic notes. Several species are confused under this
name. See also molecular data in Bohlen et al. (2011).
Pangio signicauda Britz & Maclaine, 2007
Pangio signicauda Britz & Maclaine, 2007: 26, fig. 7 (type
locality: Myanmar: Irrawaddy basin: Nanmate stream,
25�23'03"N 97�00'38"E, between Myitkina and Hopin;
holotype: USNM 378386)
Pangio superba (Roberts, 1989)
Acantopthalmus superbus Roberts, 1989: 98, fig. 74 (type
locality: Indonesia: Borneo: Kalimantan Barat: forest
stream about 1 km up Sungai Tajan from Tajan, 87 km
east of Pontianak, 0�02'S 110�07'E; holotype: MZB 3499)
